Yes, directly on the beams. I had 2133mm beams and the width was 610mm. So the tanks fit in exactly with very little space on the sides and in between the tanks.
And no I didn't have cross bracing.
I think that because yours is slightly shorter beams, with the cross bracing, yours will have less bend than mine did, if at all.
Also, I only had 1cm poly and all my tanks were fine. I'd suggest 2cm+ poly on yours.
But with all this said, my tanks were only 18in high, not 24.
Wasn't ply, was mdf. Tanks were directly on the beams/rails.